New to England, Alexander Grayson hosts a lavish party. He becomes fixated on Mina, a beautiful young woman who looks like his dead former love. Newly engaged to Mina, Harker grapples with worries over providing Mina the life she deserves.

Grayson takes Lady Jayne as a lover even as he investigates her lethal connections to The Order of the Dragon. Grayson helps Mina face a critical challenge at medical school. Van Helsing continues his quest to develop a solar vaccine to enable Grayson to conquer sunlight. Jonathan Harker strikes a devil's bargain with Grayson which has unexpected consequences.

"Goblin Merchant Men" Grayson seeks to determine whether Lady Jayne is a vampire hunter for the Order of the Dragon. Lucy mends Mina's broken heart with absinthe, romance and a whirlwind tour of Bohemian London. Grayson's machinations claim the life of Lord Laurent and earn him a powerful new enemy.

Grayson ruthlessly sets out to compromise Lady Jayne, by winning her heart. Lucy's enthusiastic plans for Mina and Harker's engagement party conceals a broken heart. Grayson's unexpected reunion with a very old friend proves a growing complication to his and VanHelsing's mission of vengeance. Van Helsing achieves a breakthrough in his efforts to enable Grayson to conquer sunlight, only to suffer an unanticipated setback.

Harker discovers that General Shaw is far from the heroic figure he presents to the public. Van Helsing's first attempt at infusing a vampiric test-subject with his solar-serum is a disaster. Lord Davenport's agents abduct and torture Renfield. Harker and Mina's engagement party sets a stage for betrayal and contretemps when a single dance between Mina and Grayson unveils their true feelings.

Browning sets out to prove once and for all that Grayson is the master vampire stalking London by hosting a board meeting outdoors in the sunlight. Grayson and Van Helsing seek to acquire the final component to make their wireless electricity technology a success to be witnessed. Meanwhile, Mina and Harker consummate their love - not knowing that Lord Davenport plots to destroy the object that Grayson loves most... Mina.

The Order tries to discredit Grayson's pioneering technology as he unveils it in a public spectacle. Browning invites Harker to join the capitalists after sensing his frustration at being unable to implicate Dracula in the General Shaw expose without revealing his own role in the fraud - but Lucy tempts the journalist with a far tastier offer. Meanwhile, the vampire tries to resist his bloodthirsty urges when he helps Mina organise a dance for the patients of her father's hospital.

Van Helsing seeks vengeance against Browning for the cuddle of his family years earlier, and Dracula struggles with his bloodlust and bid to reclaim the resonator seized by authorities on the Order's payroll. Mina faces a battle on two fronts when Lucy sets out to break her heart by seducing Harker, and Davenport plots to disfigure her after discovering she bears an uncanny resemblance to Grayson's dead wife.

Grayson, who has something he wants to get off his chest about Mina, goes to war with the Order of the Dragon just as Harker becomes a member of the group; Lucy reveals a secret that damages her friendship with Mina, who's still recovering from her attack; Browning desperately searches for his children; and Lady Jane prepares to hunt the ultimate vampire.

A public demonstration of Grayson's technology moves forward with catastrophic consequences. Lucy joins the ranks of the undead, Van Helsing exacts a terrible revenge upon Mr. Browning and Lady Jane and her huntsmen attempt to eradicate London of all vampires, culminating with a confrontation and battle-royal between her and Grayson. Finally, Mina discovers the secret of Grayson's obsessive love.