Coulson and the agents are thrust backward in time and stranded in 1931 New York City. With the all-new Zephyr set to time-jump at any moment, the team must hurry to find out exactly what happened.

With the identity of the timeline-unraveling 'thread' revealed, the team's mission to protect him at all costs leads each agent to question their own values.

A surprise leap forward in time strands Enoch in 1931 and lands the team in another unfamiliar decade; in order to stop the chronicoms from launching their future-dismantling plan, the agents must infiltrate one of S.H.I.E.L.D'.s most secure bases.

It was just another average morning on July 22, 1955, when Agent Phil Coulson realized the importance of that day in the S.H.I.E.L.D. history books. With a chip on his shoulder and a genre-bending glitch in his system, he'd set into motion a chain of events that would hopefully preserve the timeline as we know it and ensure those pesky chronicoms get the ending they deserve. What could go wrong?

After a bumpy landing in the disco decade, the team - Daniel Sousa in tow - reunites with more than one familiar face at the S.H.I.E.L.D. hangout and discovers exactly how to dismantle the Chronicoms' latest plan. But when they get too close for comfort, the Zephyr unexpectedly leaps forward again, this time to a date pivotal to not only the future of S.H.I.E.L.D. but to the future of Director Mack.

The team must scramble to rescue Mack's parents, save S.H.I.E.L.D. from a chronicom infestation and fix the ship, all before the next time-jump. Meanwhile, Daisy and Sousa find themselves at a disadvantage against a power hungry Nathaniel Malick and his goons, and Coulson will have to do the thing he does best in order to save the future.

After being unexpectedly stranded in 1982, Mack retreats to his childhood home to process the passing of his parents, while Deke gets to work scouting a chart-topping group of new agents. With no idea if or when the team is coming back for them, time seems to finally be on their side until the hugger robots show up.

With the Zephyr's time drive malfunctioning, the team hurtles toward disaster with Yo-Yo as its only hope; Yo-Yo must enlist the help of an old adversary to regain her powers.

A time storm ravages the Zephyr, propelling it toward destruction while simultaneously forcing Daisy and Coulson to relive their failed attempts to save the team.

The team arrives back in 1983, where Nathanial and Kora are hard at work building an army; Daisy is tasked with protecting Jiaying; the others quickly learn they'll need to face yet another supercharged former enemy to stop Malick.

With the help of Kora on the inside, Sibyl and Nathaniel continue their fight to shape a dark new future for S.H.I.E.L.D., managing to stay one step ahead of the agents along the way. If the team is going to turn this one around, they'll have to get creative, and maybe even a little out of this world.

With their backs against the wall and Nathaniel and Sibyl edging ever closer to eliminating S.H.I.E.L.D. from the history books, the agents must rely on their strengths to outsmart and outlast the Chronicoms.

This is their most important fight, and it will take the help of friends and teammates, past and present, to survive.