Desna and the girls struggle to keep hope alive while still working under the thumb of the new Russian regime, spearheaded by Riva. Desna remains in the dark about her gangster/gyno boyfriend, Ruval. Virginia tells Dean that she's pregnant.
Desna, now pinned under the stiletto-tipped nail of Riva's sister, Zlata, must come up with an idea to generate more customers for the clinic. Meanwhile, Quiet Ann helps Dean and Virginia get to a very important appointment, which leads to an unexpected moment of joy.
Desna's relationship with Ruval deepens when he introduces her to his mother, Matilde. Polly grows concerned when Marnie continues to pull away from her. Meanwhile, Jenn, still depressed by Bryce's recent indiscretion, falls off the wagon. After Uncle Daddy disrespects Zlata, he finds himself on the receiving end of Zlata's twisted wrath.
In an attempt to cozy up to Ruval's mother, Desna puts together a spa day for her and a couple of young girls from his charity; Quiet Ann gets a surprise visit from her brother, who convinces her to attend a family dinner.
Desna continues to grow closer to Zlata, despite the crew's increasing concern as her relationship with Dr. Ruval intensifies; Dean gets an opportunity to showcase his talents; Uncle Daddy gets his mojo back.
Faced with an agonizing dilemma, Desna and Dean take a road trip to seek guidance from a beloved childhood mentor; back in Palmetto, Jenn leans on Polly and Quiet Ann for help with a personal crisis; Virginia struggles to run the salon by herself.
Desna learns that vengeance comes at a high cost when she plots revenge on those who betrayed her; Jenn struggles to keep her feelings in check, leading her and Bryce to make a big decision that could change everything.
After a harrowing experience, Desna is offered a deal with Russian devil Zlata that's more intriguing than nail art on acrylics; Polly and Quiet Ann suffer betrayal by their boyfriends.
Polly gets an unexpected visit from her long-lost sister that sends her spinning out of control; Uncle Daddy and Roller join forces with their Dixie Mafia brethren and make a plan to take out the Russian and Haitian mafias for good.
As a hurricane bears down on Palmetto, Desna proceeds with the plans for her big day, keen for the opportunity to settle all scores; opposing forces converge at a lavish golf resort, each determined to prevail against the others.