Cresswell informs Harm that he is being relocated to London, Mac is being sent to San Diego, and they each get to bring one person from the office with them. With their respective moving dates rapidly approaching, Harm and Mac must figure out their personal lives, as well as their feelings for one another.
After being diagnosed with night blindness, Lt. Cmdr. Harmon Rabb, an F-14 Tomcat fighter pilot, is forced to change careers. Cmdr. Rabb chooses to join the Navy's Judge Advocate General (JAG) Corps. There he both defends and sometimes prosecutes Navy and Marine Corps defendants. He is sometimes assisted by fellow lawyer, Lt. Sarah "Mac" MacKenzie, and his other fellow JAG staff.
Many of Rabb's cases require him to both keep up his F-14 qualifications and travel the world, in order to bring justice to the cases he is trying.
Rabb must also deal with the fact that his father, also a former Navy pilot, never returned home after being held prisoner after he was shot down in North Vietnam. Rabb believing his father is still alive continues the search he has pursued to find his father, since his teens.