Now that Wilhelmina has reestablished herself at MODE, Daniel must make some changes due to her manipulations. Meanwhile, Betty makes an important decision regarding her life, and Ignacio gets a job at a fast food restaurant.
Wilhelmina, now preparing for her first issue as editor, puts Betty in a difficult situation when she offers her a job as her assistant. Meanwhile, Christina becomes suspicious of Stuart's true intentions, Daniel faces a custody battle to keep Daniel Jr. in the United States, and Hilda's romance with Coach Diaz reaches a new level of hotness.
Everyone at Mode is stunned when an accident occurs. Later on, Betty tries to figure out what really happened when a detective comes by with the same purpose. Meanwhile, Ignacio is shocked when he learns a truth about Hilda and Coach Diaz.
Kimmie asks Betty for help when she finds herself in need of a job. Meanwhile, Justin auditions for a Broadway musical, and Daniel signs up for a matchmaking service but finds a surprising result with regards to his match.
Betty has a competition with Kimmie. Ignacio decides to vote for the first time ever. Hilda takes advantage of the election traffic to be able to promote her shop. Meanwhile, Wilhelmina tries to hire a new financial officer whom Daniel hates.
Wanting to grasp the attention of her neighbor Jesse, Betty decides to work overtime. Meanwhile, Amanda becomes Betty's roommate when the former finds herself in financial instability, and Cliff proposes that he and Marc move in together.
Betty tries to prevent an issue of Mode from shipping because it can result in a PR mess for the magazine. Meanwhile, Hilda's business license is denied by the city, forcing her to go to other means to keep her salon alive.
Marc and Betty compete against each other for the Young Editors Training Program but only one spot is available. Daniel learns he is drawn towards Molly. Wilhelmina attemps to get Connor to notice her.
Amanda and Betty are forced to work together for an article called "How I Blew Ten Grand Without Spending a Dime."
Betty finds it hard to impress her idol, Jodie Papadakis who is an editor at YETI, while making a mess of her work and family life. Betty's has a disastrous encounter with the assistant at ELLE magazine.
Betty finds herself stuck between work and home life as she tries her best to appease both Hilda; Daniel, Molly, Connor and Wilhelmina.
Betty comes up with a solution to deal with her personal problems but she has her sexy neighbour Jesse who keeps her distracted from her goals. Daniel does his best to cover his tracks as Connor is adament on unveiling Molly's admirer.
It's Claire's 60th birthday and Betty makes her a video which reveals some unintentional footage of Mrs Meade. Also, Daniel and Wilhelmina are finding it hard having someone new in their lives.
Fashion Week hits town which causes problems for the staff at Mode. Betty has to split her time between working on her Yeti assignment and working with a new designer. Elsewhere, Wilhelmina has lost her way because of Connor and there's a new person working at Mode.
Betty and Marc team up together to analyse Mode's budget. Matt continues with his pursuit to win Betty while she deals with some startling news on Facebook by a former flame.
The Suarez family attempt to buy their rented house from their landlord. Ignacio goes head to head with Frankie Burrata, a celebrity chef. While everyone in Mode comes to deal with their shock.
Betty comes face to face with Matt's mother. Wilhelmina devises a way to make money fast whilst Daniel deals with Molly. The Suarez family attempt to set Hilda and Archie up on a date.
Daniel and Wilhelmina make Mode's issue based on sex to spice things up. Matt is hesitant to take the next step with Betty. Marc and Amanda provide Betty with a makeover.
The Suarez family meet Calvin Hartley, the billionaire father of Matt's. Betty aims to get personal with Matt while Daniel tries to save Mode.
Baby William is the unlikely hold up in Mode receiving the money it needs to save the company, whilst Hilda is asked to help with a political campaign.
Betty, Matt and Marc come together in order to succeded in their final YETI project, a photo session with singer, Adele. Love is in the air for one lucky couple as proposals are made. Claire and Wilhelmina continue to argue.
Henry forces Betty to acknowledge she still loves her former fiance. Daniel wins the media over with his transformation as a devoted husband. While Wilhelmina tries to get information from Victoria Hartley about the hold Claire Meade has on Cal Hartley.
In the season finale Marc and Betty are left to battle it out for the same job after the passing of a Mode editor. Wilhelmina threatens Claire to resign or her dark secret will be shown to Cal. Matt struggles with his jealousy, while Justin nervously waits to see if he has a place in the performing arts school.