Cassey deals with living the single life now following Cappie's rejection, while Cappie learns about Max's leaving. Elsewhere, Rusty risks losing his honors program if he can't raise his organic chemistry grade.
It's father-daughter weekend at ZBZ, as tension builds at the house after the secret of whom Rebecca's kissed is revealed. Meanwhile, Rusty tries to impress Jordan's father with sports, and Dale and Cappie explore different religions.
As Rusty and Jordan are awkwardly deciding when to take their relationship to the next level, the Gotcha Game comes between them as the final two players left in the game. Leading Cappie to become Rusty's bodyguard and Rebecca as Jordan's, as the happy couple battle it out so their house can win the game who will pull the trigger first. Meanwhile, Casey takes over as the ZBZ Panhellenic delegate, butting heads with the president, Katherine, over the annual "Undie Run". After Katherine has the event canceled, Casey uses Evan's help to get enough signatures to overturn the decision and save the traditional event by using it as a fundraising event for charity. And Calvin and Grant are trying desperately to find some alone time so they can be together away from their raucous Omega Chi brothers. In the end, they decide to leave campus for a romantic dinner, but are surprised by who turns out to be their waiter.
It's Dry Weekend at Cyprus Rhodes University, and Casey plans a non-alcohol party night amongst all of the houses but, when she and Evan are partnered together to patrol each of the houses for signs of any prohibited beverages, they each have a brownie at the Psi Phi house unaware of its secret ingredient, causing them to hide out in the ZBZ bathroom until the effects wear off. Meanwhile, Ashleigh frantically tries to find Casey, and Fisher offers to help, making her further question whether or not she should take him back. Also Grant's high school girlfriend, Whitney surprises Grant andd Calvin by visiting for his birthday. So Calvin enlists Rebecca's help on a double date as he patiently waits for Grant to tell Whitney the truth about himself. And Rusty learns that he earned a new nickname in the engineering school because of his slipping grades so, he decides to approach the infamous Dr. Larsen to see if he will sponsor Rusty in an experiment and, Cappie joins him for moral support and attempts to flirt with the engineering girls.
Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers. Meanwhile, dissatisfied with her college experience, Jordan wonders how to make her time at CRU more fulfilling and, in an effort to prove his love and commitment for Jordan Rusty decides to lavaliere her. When Rusty divulges his plans to his KT brothers, they warn him of the Kappa Tau curse that occurs when a brother lavalieres his girlfriend. Elsewhere, Cappie helps Evan find a part time job to help his financial situation, but when he meets one of Evan’s new and very attractive co-workers, Lana, he decides to get a job too.
It's Founders Weekend at the ZBZ House and Casey uses the opportunity to scrap for donations from the alums while Rusty tries to keep Jordan from dropping out of school.
Casey's detective skills inadvertantly start a brawl between the Kappa Tau's and the Omega Chi's. Meanwhile, Rusty, Calvin, and Dale take a trip to Comic-Con as Ashleigh and Fisher resume their relationship but still have a few issues.
Rusty thinks Cappie might be ruining his new relationship with Lana, so he convinces Dale to help him save Cappie from himself. Casey and Ashleigh try to do a mixer with ZBZ and Omega Chi to get the sorority its reputation back.
Everyone is packing their bags and heading home for Thanksgiving, but Casey and Rusty find themselves stranded at CRU when their parents leave them behind for Maui. Will Thanksgiving at CRU beat going home for the holidays?
Will Cappie's and Casey finally get back together? And if so, what does this mean for Cappie and Evan's friendship. Will Ashley be supportive? As one of the biggest blue ribbon competitions Songfest is approaching, the ZBZs are determined to win it all.
The ZBZ ladies try to cover their tracks after the Gamma Psi house burns down; Rusty has difficulties in his position as pledge educator; Casey plans a Fairytale Ball to raise funds.
The ZBZ's hear that their candle is responsible for the Gamma Psi fire, and Casey cracks under the guilt, putting the house in danger. Hoping to cheer up the house and boost her marketing major, Ashleigh enlists Rusty and Dale in a philanthropic "Geek Auction", but Casey is the one who has to play dirty to save the house. Can Casey save the day?
It's an All-Greek baseball game and with the hope of winning more blue ribbon points, the ZBZs partner with the Omega Chi's, puting Capie and Casey in opposite teams. Meanwhile, Rusty is not doing so well as pledge educator of the KTs and everything gets worse when Calvin asks him to spend quality time, getting to know Grant, who previously duct-taped him to a pole. Meanwhile, Evan and Rebecca become more serious but because Rebecca doesn't believe in love, Evan turns to an unlikely source to find the meaning of love.
Casey and a resistant Cappie, try to talk about the future after the "G" word Graduation. The situation gets worse when a handsome Washington politician, encourages Casey to view politics as a career, after she helps him with his voter registration campaign. Is Cappie ready to grow up? Meanwhile, Rusty is selected to appear in the cover of EW(Engineering Weekly), but an argument with Dana leads Rusty to question himself and his project just before the interview.
It's Valentine's Day! Casey and Cappie compete over who can come up with the most romantic gift. Also, Casey sets Katherine up with Rusty in an attempt to reconcile with her. Ashleigh is worried that new boyfriend Pete will turn out to be another Fisher, especially after Natalie implies that she and Pete are in a relationship. Evan attempts to bring out Rebecca's obscure romantic side with a special gift. Meanwhile, Calvin is worried that he and Grant are losing their spark, mainly after observing Grant's new beauty ritual.
The Hilgendorfs have moved next door to the Kappa Tau house, and their presence is putting a damper on the KTs' festivities. When their interferes too much with the fraternity, Rusty and Cappie decide to to bring Professor Tom Hilgendorf in and make him a pledge. But once inducted, Tom doesn't seem to want to leave, leading Cappie to rethink his plans on graduating.
Everyone at Cyprus Rhodes is preparing for Mardi Gras blowouts, except for Casey, who must prepare for the LSAT. While she continues to push Cappie to declare a major and make plans for the future, he is too busy announcing the next parade at the Kappa Tau Mardi Gras party. Meanwhile, Joel offers Casey some advice and support for her upcoming exam.
Casey and Cappie get a surprise visit from Cappie's parents, who invite them to a camping trip. While reluctant Casey agrees to go, in hope that they might encourage Cappie to choose a major and graduate with her. But she quickly learns that they are free-loving hippies who think Cappie should just relax and enjoy his college experience.
A golf tournament will determine the top fraternity on campus; Katherine agrees to bring back the Greek Goddesses Pageant; Calvin and Grant feud over where to spend spring break.
The gang returns to Myrtle Beach for spring break; seniors consider their futures.