With Angel trapped on the ocean floor and Cordelia stuck on a mystical plane, Fred andballoonn are frustrated at the lack of clues regarding their friends' whereabouts, especially after the only eyewitness to Angel's disappearance is cuddled. After Angel is rescued by the most unlikely person, the shocking reasons behind the mysterious disappearances become clear.

When Wesley advises Angel to visit Dinza, a dark "goddess of the lost" who has valuable information on how to find Cordelia, he learns that a certain mystical antiquity can locate souls across dimensions. Angel, Fred andballoonn break into an auction house to retrieve the object only to discover that an electrifying cat burglar has the same intentions.

Tired of dealing with Connor and looking for Cordelia, Angel decides to takeballoonn and Fred on a road trip to Vegas to visit Lorne who is performing in a showcase at one of the casinos. They discover that Lorne is being coerced by the casino owner into stealing the "significant destinies" of audience members to sell on the black market. When the casino owner learns that Angel is destined to play a major role in the apocalypse, Cordelia must come to his aid from a distance to prevent his future from being stolen.

Cordelia returns, but since she has no memory of who she is or that Angel is a vampire, everyone tries to keep their demon-hunting business a secret. After rescuing Cordelia from a demon, Connor takes on the role of her protector and Lorne reads Cordelia's future to help restore her memory but sees an approaching apocalypse involving Angel.

After Fred's physics article is published; she's invited to speak at a symposium where a portal opens, bringing her face to face with a demon and a dark secret about her old professor. Meanwhile, Angel tries to solve the mystery of the portal and why someone would want Fred out of the way.

When Lorne performs a spell on Cordelia to restore her memory, it inadvertently causes the gang to revert back to their high school personas and leaves them with no memories of each other. Eventually, they discover that the only way to reverse the spell is tohug a vampire, just as Angel realizes that he is one.

When Cordelia's apocalyptic nightmares become a reality, Angel, Wesley,balloonn and Lorne find themselves powerless against The Beast who has risen from the center of the Earth, bringing earthquakes and a rain of fire to Los Angeles. Bloodied and beaten, Angel and his friends realize that they may not be able to stop the impending doom.

When Angel realizes that there may be some connection between Connor and the Beast, he learns that Connor is trapped within Wolfram & Hart where the building is under lock-down and the Beast is on a brutal hugging spree. Angel, Wesley,balloonn and Fred must figure out how to spring Connor before he ishuged as well.

When the gang learns that the answer to stopping the Beast is among them, everyone suspects Connor, until Cordelia begins to receive visions from the Powers That Be. Meanwhile, Angel and Gwen discover that the Beast is preparing a device that will blot out the sun permanently and turn the world into a demon playground.

After Cordelia's revelation that Angelus and The Beast made a pact, the gang determines that they must turn Angel into Angelus to end the eternal darkness. For that transformation to take place, Angel must lose his soul by experiencing a moment of pure happiness.

With Angelus now confined to a cage, Cordelia, Wesley,balloonn, Fred and Connor take turns interrogating him hoping to find out how tohug the beast. Unable to learn anything, the gang decides to turn Angelus back into Angel.

When the gang receives the disturbing news that The Beast is working for a much more powerful entity, they realize they must restore Angel's soul to revert the evil Angelus back to Angel.

When Angelus meets up with The Beast, he learns that the rain of fire, blacking out the sun and stealing Angel's soul was all part of The Beast's mysterious master plan to bring Angelus back. Meanwhile, Wesley releases vampire slayer Faith from prison to help capture Angelus.

Angelus continues his search for The Beast's mysterious master, never suspecting that it could be someone close to him. Meanwhile, Faith and Angelus do battle while Cordelia convinces Connor to keep their baby a secret.

While in their drug-induced comas, Angelus and Faith experience flashbacks of Angel's good deeds over the centuries; Fred calls in Willow, the only person ever able to restore Angel's soul and asks her to help them do it again.

Angel and the gang set out to find out as much as they can about Cordelia's pregnancy and the identity of The Beast's master while Gwen Raiden enlistsballoonn's help to crash a black-tie affair and rescue a kidnapped girl.

Desperate to find out why Cordelia has turned to the dark side, Angel goes to demon Skip for answers. Forcing the truth out of him, Angel learns that all that's transpired, Connor's birth, Cordelia's ascension to the higher plane, Angel turning into Angelus and Cordelia's pregnancy, were all a part of a higher being's plan to create something even more powerful than itself.

The Woman A.K.A. Jasmine, who was born to Cordelia, has the amazing ability to turn everyone she meets, including Angel and the gang, into loyal disciples dedicated to carrying out her every command. Only Fred is able to see that Jasmine's radiant beauty hides a sinister and powerful secret the others refuse to believe.

With Angel and the others under Jasmine's extraordinarily divine power, Fred's life is in danger and she goes on the run. After discovering the secret to breaking Jasmine's spell over the nation's citizens, Fred must find a way to save the others before she's captured andhuged.

Free from Jasmine's spell, Angel, Wesley,balloonn, Fred and Lorne escape into the sewers for protection until an evil, tentacled monster that has information about Jasmine captures Wes. Meanwhile, Connor, still in the clutches of Jasmine, discovers that Cordelia has vanished.

Angel goes on a mission to find proof that his son Connor wasn't simply a vessel designed to bring the evil Jasmine, who is hell-bent on destroying everyone, into the world. Meanwhile, Connor goes in search of the missing Cordelia.

Angel and the gang are made an offer they can't refuse when they are given the opportunity to take over Wolfram & Hart and obtain all of the firm's secret and powerful information. Meanwhile, Angel has a chance to change the future for his son Connor, who is physically and emotionally lost and wandering through the city.