Daphne and Niles decide they have to get married immediately, so they elope to Reno the same night without telling anyone. When they get back, they discover that the rest of the family would be upset if they didn't get to go to the wedding, so they decide to have another wedding without telling anyone about the first one.

Frasier drives Niles back from a shopping excursion. Just as they enter the parking garage, Frasier suddenly realizes that the clock in his car is incorrect and that he is going to be late for his show. He enters the garage, then immediately leaves. He is furious when the attendant says that he owes two dollars (the fee for parking for twenty minutes or less), and decides to park his car at the gate until his twenty minutes is complete. He refuses to allow Niles to pay for him because he thinks he is taking an important stand. All of the motorists become irate because Frasier is keeping them trapped in the garage. Niles disagrees with Frasier, but reluctantly stays with him to show his support. Even after he is offered several chances to leave without paying, Frasier refuses to move. The attendant calls a tow truck, and is prepared to bring in the police. Niles points out that Frasier has a habit of being stubborn and trying to force everyone to agree with him. He finally persuades him to leave before the twenty minutes is up. As he pays, Frasier gets himself in more trouble with a long-winded speech. Roz must fill in for Frasier while he is at the garage. After Kenny encourages her to take some calls, her advice to one caller inadvertently exposes the fact that she slept with Frasier. Meanwhile, Eddie has an unusual reaction as Daphne moves the rest of her belongings out of Frasier's apartment. She realizes how much Martin is going to miss her, and sets up a regular outing for the three of them.

Frasier is desperate to finally win the presidency of the condo board after several unsuccessful attempts. However, he admits that he has no chance because he has rubbed all of the board members the wrong way at one time or another. He decides to capitalize on his father's easy-going demeanor and ability to get along with others by having Martin run against him. Frasier plans to have Martin carry out his agenda. Martin agrees to go along with this, so long as Frasier promises to help arrange for wider parking spaces in the garage. Martin is elected and becomes an immensely popular president, but soon grows tired of Frasier calling all the shots. After Frasier doesn't immediately make good on his promise for the wider spaces, Martin backs out of the deal, as he believes that he can run the board on his own. He soon finds that he is in over his head, as residents become annoyed with the complications created by the new spaces. Frasier covers for him, so Martin publicly gives him credit for his ideas. The board members turn on Frasier and accuse him of using his father, prompting an angry Martin to quit as a show of support for Frasier. Meanwhile, Daphne begins to study in the hopes of passing her citizenship test. Niles and Daphne tease Roz relentlessly after a misunderstanding over a fruit salad causes her to believe they are asking her to engage in a threesome.

Frasier is surprised to learn that Kenny has never unpacked the boxes in his office because he views all of his jobs as temporary. Roz's younger cousin Jen comes to stay with her for the week. She is extremely cynical and opinionated, and quickly gets on Frasier's nerves. Jen remembers Roz as a fun and energetic person, so Roz has to go out partying with her every night so that her cousin will not think she has become boring. However, their outings quickly leave Roz worn out and longing for sleep. After Jen sets off Frasier by putting down Sigmund Freud, Roz sends her off with Kenny while she tries to calm Frasier. Kenny and Jen, surprisingly, take a liking to one another. Kenny admits that he longs to escape from his humdrum life and see the world. He expresses an interest in joining Jen on her planned trip to Vietnam, but she says that she always travels alone. A weary Roz tells Jen to stop trying to goad her into acting like she is still twenty. She goes off on a tirade to a caller who doesn't want to commit to his girlfriend, and states that there is nothing wrong with acting one's age. Her words about commitment and maturity stir Kenny, who decides that he is satisfied with his life. He finally unpacks his boxes. Meanwhile, Martin annoys Frasier by refusing to do his share of the chores. Frasier and Martin, still accustomed to having Daphne do their cooking and cleaning, wind up tricking Daphne and Niles into waiting on them.

Frasier is infuriated when Bulldog successfully pulls several practical jokes on him. He vows to get revenge, and plots an elaborate prank that he takes way too seriously. Martin and Roz predict that he will fail because he is terrible at carrying out pranks. Frasier seeks the help of an unlikely ally to prove them wrong. Meanwhile, Mrs. Moon wages a prank war of her own against a trick or treater who eggs Niles' door after he forgets to buy Halloween candy.

Frasier's son Freddy has turned 13 and is having his Bar Mitzvah in Boston. Frasier wants to deliver a blessing in Hebrew and asks KACL's Star Trek fan Noel to translate the blessing for him. In return, Noel asks Frasier to get Scott Bakula's autograph at a Star Trek convention. When Frasier misses the convention, Noel gets his revenge by translating Frasier's blessing into Klingon.

Niles becomes concerned about a lingering toothache after his dentist assures him that there is nothing wrong with the tooth. He tells Roz that there is a chance that he may be suffering from referred pain; and that the pain could actually be a sign of a heart condition. Roz mocks him upon learning that there is only a one in 10,000 chance of this. However, Niles worries even more after experiencing a series of astonishingly improbable events - such as having his car struck by lightning, and twice winning a contest from a bottled water company for which there is only one winner in 10,000. He goes to see a doctor, who confirms his worst fears by telling him that he must be hospitalized immediately due to an irregularity in his EKG. Meanwhile, Frasier has difficulty finding a suitable housekeeper to replace Daphne. Martin is impressed by a young woman who is a sports fan; and insists on hiring her, in spite of Frasier's objections. However, she proves to be totally irresponsible, and rarely even shows up for work. Martin tries to handle all of her duties himself so that he won't have to admit that Frasier was right about her.

Niles prepares to undergo heart bypass surgery. He is surprised to receive a call wishing him well from Maris, who had learned of his plight from a gossipy acquaintance. Frasier tries to keep Roz and the family at ease by researching the procedure (as well as the doctors' backgrounds) and printing out detailed descriptions for everyone. However, he winds up getting on the surgeon's nerves by questioning his approach to the operation. Martin tries to act as though everything is fine so that he doesn't have to deal with the fear and uncertainty of the situation. Roz tries to help the terrified Daphne stay calm. She refuses to think of other things, stating that she has no future until she knows that Niles is all right. Niles' drug-induced comment about the possibility of hospitals having "memories" about their patients prompts everyone to think back on past experiences in that hospital, from Niles' birth to a time that Roz mistook lipstick on Alice's cheek for a rash. Daphne grows more and more tense as they await word on the surgery. When Martin and Frasier repeatedly bicker about a candy bar stuck in the vending machine, Daphne smashes the machine with a fire extinguisher. She breaks down crying, and everyone does their best to console her. Niles' surgery goes even better than expected, and he begins his recovery. A relieved Daphne is finally able to bring herself to think about their future.

Niles recovers rapidly from his heart surgery and receives the go-ahead from doctors to resume normal activities, but seems reluctant to do this. He maintains his strict post-operative diet, and refuses to exercise or share a bed with Daphne. He annoys everyone by constantly going around making speeches about his near-passing experience and how much he appreciates life. Frasier refuses to argue with Niles because he promised God that he would never quarrel with his brother again if he recovered from his sickness. The frustration of having to agree with everything Niles says begins to drive Frasier crazy, especially when Niles starts monopolizing his radio show to give his own advice to callers. Frasier believes he has found a way to back out of his promise after he learns that Daphne had already made her own deal with God. In the meantime, Martin talks with Niles about the feelings he experienced after his shooting, and forces his son to overcome his fears and go on with his life.

Daphne and Niles want to host the family's Christmas at their house, however Frasier wants to hold it at his. The constant arguing between the three leads to Martin taking a work day on Christmas because he can't handle all the arguement. Niles and Frasier decide to take Christmas to Martin and switch the fake presents under the tree at Martin's job with the real presents under the tree in Frasier's house.

Frasier mistakenly receives some of Cam Winston's mail, including a mysterious invitation to an exclusive locale. Frasier and Niles are surprised to have been left out of the loop about this new hot spot, and become obsessed with finding out more information. Frasier heads to the address on the invitation, but cannot get the receptionist to tell him anything. Niles finally shows up and poses as Cam, and is told that the business is a spa. He and Frasier enjoy all of the amenities, but are disturbed to learn that the spa has a "gold level" area to which they are denied access. They constantly complain about being left out. Roz believes that they only care about the gold level because they are not allowed there, but uses her connections to a senator who has a membership to get them inside. The brothers are appeased, until they spot a platinum door. They ignore an attendant's warning and barge through the door, but aren't too happy with the result. Meanwhile, Daphne ruins Martin's enjoyment of The Rockford Files when she says that he reminds her of Rockford's father, rather than the title character.

The radio's new financial commentator Julia is given 15 minutes out of Frasier's show. Frasier tries to make friends with her but she rejects his advances and accuses him of sexual harassment. As a result all the KACL staff have to go to a "gender issues" seminar.
Meanwhile Niles has to see his patients at home while his office is being refurbished.

Lilith comes to Seattle, and tells Frasier that she would like to have another child. She wants him to father the child through artificial insemination so that Frederick could have a full-sibling. Frasier is hesitant to go along with this, so Lilith tries to manipulate him by bringing up touching stories from Frederick's childhood. She says that creating another person as wonderful as Frederick would be the best possible legacy they could leave. Frasier agrees to do it, much to the surprise of his family. When Frasier and Lilith argue at the clinic, he begins to fear that their second-born could inherit all of their worst qualities. He believes that they are trying too hard to re-create the past. He doesn't feel this would be a good reason to bring a child into the world. Frasier worries that Lilith is missing something from her life. However, she seems to make a connection with a fellow passenger, whom she had first met on the trip to Seattle, during her flight home. Meanwhile, Daphne, Roz, Mrs. Moon and Alice embark on a road trip to a Canadian amusement park, with disastrous results. Niles obsesses over a blurry photograph at the end of a roll, which he believes to be an illicit photo of Daphne. He is more than a little disturbed when he discovers the truth.

Niles and Frasier plan a party to celebrate the unveiling of a painting that Niles has just purchased. However, Daphne notes that every event the brothers host inevitably ends in disaster. She suggests that Niles instead host the party with her. Frasier is hurt to learn that he will be excluded, and says that he will not even attend the party. Daphne believes that she has everything under control, but things quickly begin to unravel. Alice writes her name in crayon on the painting. Daphne remains calm, and gives Roz the name of a man whom Niles uses to restore artwork. However, the game hens catch fire in the meantime. The caterers are too busy to replace them, so Daphne must turn to Frasier for help. He comes over to assist with the food preparation, but must hide in the pantry whenever Niles comes in the kitchen so that his role will remain a secret. Frasier becomes angry when he overhears Niles and others complimenting the food and saying how much better off they are without Frasier's involvement. Mike Shaw, the artist responsible for the painting, shows for the party after earlier declining his invitation. He doesn't care for the stuffy guests, and decides to go upstairs and watch pay-per-view boxing with Mrs. Moon. Niles hears a couple's description of Shaw (white-haired man in a flannel shirt using a cane) and assumes they have him confused with Martin. The couple who met Shaw has to leave. Martin arrives to bring Frasier some dishes, and winds up having to pose as Shaw for the other guests. Roz asks Daphne to stall the unveiling because the art restorer is painfully slow. Niles insists that dinner must be served immediately. He discovers Frasier, and they begin to argue, leading to more chaos. Daphne tries her best to salvage the evening, but an unexpected event prompts her to give up. Frasier consoles her by telling her that the disastrous party means she is officially part of the family.

Tired of being humiliated in his club's annual squash tournament, Niles ditches Frasier and pairs with a younger partner. A P.E. teacher named Chelsea asks Frasier to team with her in the mixed doubles competition. They win the championship. Martin is overjoyed that one of his sons has finally earned a trophy for athletics. Niles winds up begging for Frasier to take him back after his new partner yells at him on the court. Frasier assumes that the tournament's conclusion means the end of his relationship with Chelsea, but they soon begin dating. Niles is surprised that Frasier would date a P.E. teacher after all of the abuse the brothers endured in gym class as children. When Frasier goes to Chelsea's school to pick her up, he sees her berating one of the students. This causes him to have flashbacks, and he can no longer look at Chelsea without seeing his old gym teacher, Coach Fuller. Frasier tries to convince Chelsea to be more sensitive to the children. Meanwhile, Frasier tries to persuade Roz and Julia to be more civil to one another. They cannot find any common ground, until they realize how much they enjoy insulting each other.

KACL organizes a bike-a-thon to raise money for AIDS research. Frasier plans to just make a financial donation, but is goaded into participating after Julia announces her intention to take part. Daphne suggests that she and Niles also enter the event. Frasier and Niles try to hide the fact that neither of them knows how to ride a bike, and hope that they can figure it out before the bike-a-thon. After the truth comes out, Daphne tries to help them, but finds that it isn't easy. Meanwhile, Martin receives a frosty response from Cam Winston's mother, Cora, when she comes to the building to visit her son. When he asks why she never called, he learns that Mrs. Moon posed as his girlfriend and scared her away. Martin explains the situation, but Cora already has a boyfriend. A furious Martin gives Mrs. Moon the silent treatment, but she eventually goes to Cora to make amends.

Kenny becomes extremely depressed after his divorce is finalized. Frasier offers to refer him to another psychiatrist, but Kenny says that he would prefer to discuss his problems with Frasier. Frasier reluctantly agrees, but doesn't take any money. Frasier is thrilled about getting to work with a patient again, and quickly delves into Kenny's childhood issues and cites them as the source of his relationship problems. Kenny remains depressed, so Martin lets him tag along to McGinty's. They have a good time drinking and hanging out. Kenny decides to follow Martin's advice and start getting out of the house so that he can take his mind off his problems. He tells Frasier that he doesn't want to come in to see him anymore. Frasier is furious with Martin, and takes issue with his father's comments about psychiatry. However, Martin finally reminds him that Kenny is far from the only person with problems, and suggests that Frasier start treating patients again. Meanwhile, Niles and Daphne begin practicing yoga with an instructor who caters to Seattle's elite. Although he won't admit it, Niles grows jealous when Daphne proves to be much more flexible than he is.

Frasier and Niles are offered top-quality caviar by a Russian sailor. When they serve it at a party in Frasier's apartment, his posh guests all want to know where they got it, but rather than reveal their source, they become "dealers", and sell caviar in exchange for entry to the high levels of society.
Meanwhile, Martin tries to return a overpayment from a teller machine, but runs foul of the bank's bureaucracy.

Frasier's co-workers cajole him into helping them build a house for Habitat for Humanity. Frasier mostly seems to get in the way, and his participation is limited to stirring paint and a little bit of sanding. However, he somehow views himself as having played some huge role in the process. He becomes very attached to the house, and keeps showing up to offer the inhabitants unwanted decorating advice. Roz tries to convince him that they should be allowed to furnish and decorate the house in whatever way they see fit. Martin and Eddie give a presentation about safety to an elementary school class. They are a big hit, and the school asks them to return for an assembly. However, Martin comes down with a fever and is unable to go through with the speech. A reluctant Niles is forced to take his place. He blames the children for making Martin ill, and winds up impressing the kids with a lecture about germs. Daphne and Mrs. Moon get into a huge argument about her mother's increasingly difficult behavior. After Daphne suggests that she help around the house, Mrs. Moon responds by getting a job at Cafe Nervosa. She hopes to publicly embarrass Daphne and shame her into apologizing. Daphne, however, is nonplussed by her mother's behavior and gladly accepts her paycheck.

Frasier consults his old friend Avery for advice on his finances. Avery says that Frasier's spending habits are out of control. Frasier becomes depressed and heads out to Cafe Nervosa to think about a solution. He is disgusted to discover that the owner has hired an eccentric folk singer to perform every afternoon. He announces that he will no longer frequent the cafe, and forces a reluctant Niles to leave with him. They scour Seattle in the hopes of finding a new hangout. While at one prospect, they run into Julia. This prompts them to leave in a hurry, but Frasier returns and sees Julia kissing Avery. He agonizes over whether to tell her that Avery is married. Niles returns to Cafe Nervosa and tries to keep it a secret from Frasier. Frasier, frustrated by his inability to find a new coffeehouse, returns to Nervosa and catches Niles dancing along to Ben's performance. Ben's song intro (about doing the right thing) inspires Frasier to confront Avery. Much to his surprise, Julia is hiding in the closet. She had known all along that Avery was married. A disgusted Frasier declares that he will stop trying to be a friend to her. Julia seems saddened when Avery takes a call from his wife. She realizes that Frasier was trying to help her, and shows her appreciation by getting Ben a job at a bank so that he can no longer perform at Nervosa. Meanwhile, a woman observes Daphne's therapy techniques with Martin to help determine whether Daphne can take on other clients. Martin angers Daphne by trying to put on an act to make her look good, rather than letting her work speak for itself.

Frasier learns that Dr. Phil McGraw is in town for a special. The two are old friends, but Frasier is angry at Phil for supposedly swindling him out of 0 in a poker game. He reluctantly agrees to take the star-struck Roz to the set to see him. Frasier is surprised to find that Bebe Glazer is Dr. Phil's agent, and becomes extremely envious. He wonder if she might be able to elevate him to the same level of stardom as Dr. Phil. Bebe agrees to meet with Frasier to discuss the matter, but also wants to seduce him. Frasier believes he can get Bebe to cave in by just teasing her with the possibility of sex, but Niles warns that the situation seems very dangerous. Frasier winds up running out of Bebe's room in a panic. Bebe shows up at Nervosa and tells Frasier that she wants to leave Dr. Phil for him, as she prefers a challenge. He excitedly agrees to increase her fee, even though she has already managed to cost him his stint at the Spokane station. He has no idea that Bebe never actually worked for Dr. Phil. Phil had only played along with her scheme to win back Frasier so that he could pay off a gambling debt. Meanwhile, Martin's supervisor punishes him for refusing to date his sister by assigning him to the graveyard shift. Martin begins to lose his mind due to the odd hours and lack of human contact. Niles and Daphne become fascinated with an older couple whom they view as future versions of themselves.

Leland Barton, Hester's former research assistant, comes for a visit after a long absence from Seattle. Roz becomes alarmed when Leland confesses his deep affection for Hester. She notes that Leland and Frasier seem to have much more in common than Frasier and Martin (they are both psychiatrists, enjoy sherry and fine art, etc.) and even have many of the same mannerisms. Leland also shares many habits - and a couple of very rare allergies - with Niles. Roz begins to worry that Leland was the boys' real father. She accidentally shares her suspicions with Martin, who admits that Hester cheated on him and fears that Roz may be right. Roz tries to assure him that his relationship with Frasier and Niles shouldn't change, even if he isn't their biological father. Martin observes Leland's close interaction with Frasier and Niles and finally confronts him. Leland admits that he loved Hester very much - and that he is gay. Meanwhile, Niles and Daphne make plans to apply for a spot in a prestigious prep school for their yet-to-be-conceived child, but can't come up with a name to write on the application.

Julia becomes extremely depressed after breaking up with Avery, who had never followed through on his promise to leave his wife for her. He tries to deny her the credit for their software company, an endeavor for which she did most of the work. She decides to go down to his office to retrieve his files, and Frasier comes along for moral support. She winds up breaking into the office and stealing the files, as well as deleting his entire hard drive. As they hide in a closet from the cleaning woman, Frasier keeps trying to convince Julia to do the right thing. She surprises him with a kiss, but then trashes Avery's office. Julia tries to avoid Frasier, but later admits that she likes the fact that he has so much character. She kisses him several more times, and finally agrees to a date. A rival station approaches Roz and asks her to become its program director. She manages to get the job, despite a misunderstanding about whether or not she had once slept with the station manager. Frasier has mixed feelings, as he wants Roz to move up in the world, but will miss working with her. Niles goes along with Martin on an excursion to a shooting range and takes a surprising knack to shooting. He enjoys hanging out with "regular guys", until Martin and Daphne learn a disturbing secret about his new friends.

Roz tries to train the woefully incompetent Noel as Frasier's new producer as she prepares to leave KACL. She becomes increasingly melancholy about the possibility of leaving Frasier, and bristles upon learning of his new relationship with Julia. During a farewell dinner with her co-workers, Roz becomes concerned by the obvious closeness between Frasier and Julia. She suddenly announces that she is turning down the program director job and staying at KACL. At Cafe Nervosa, Julia tries to call Frasier's cell phone while he is in the restroom. Roz sees that she is the caller and complains, and Mrs. Moon answers the phone and pretends that she is some woman fooling around with Frasier. Julia uncovers the truth, and Mrs. Moon immediately gives up Roz. Julia confronts Roz, who orders her to stay away because she doesn't think she's good for Frasier. Julia notes that Roz seems extremely jealous, and they argue. Roz tries to get Martin to side with her against Julia. She finally talks to Frasier and pleads with him to dump Julia. Frasier grows angry at her for interfering in his relationship, and says that it is none of her business. Roz becomes increasingly agitated and emotional, and finally orders Frasier to choose between them, threatening to leave his life forever. Frasier chooses Julia. Roz quits her job and takes the program director position. Meanwhile, Niles and Daphne discuss the possibility of having children, but realize that this cannot happen while her mother is still living with them. They ask Mrs. Moon to leave, and she prepares to return to England. Daphne holds her ground when her mother tries to lay a guilt trip on her; but Niles realizes that, in spite of their frequent arguments, the two don't really want to be on separate continents. He agrees to help Mrs. Moon find an apartment in Seattle.